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We introduce Symbiosis, a framework for key-value storage systems that dynamically configures application and kernel cache sizes to improve performance. We integrate Symbiosis into three production systems — LevelDB, WiredTiger, and RocksDB — and, through a series of experiments on various read-heavy workloads and environments, show that Symbiosis improves performance by 1.5× on average and over 5× at best compared to static configurations, across a wide range of synthetic and real-world workloads.